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1. 10. Plastids Chloroplast Amyloplast Leucoplast Chromoplast
(from greek plastikos: “molded”) Chloroplast Amyloplast Leucoplast Chromoplast
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Rapid Changes in plastid shape
Fig 1-43 Light micrograph of plastids with thin, tubular stromule extensions
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All plastids are developmentally related to proplastids
Plastid developmental cycle and the inter-conversion of various plastid types
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Fig. 1-45. TEM showing a proplastid (left) adjacent to a mitochondrion
in a bean root cell
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TEM illustrating phytoferritin deposits inside a proplastid
in a root apical meristem cell of soybean
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2) Amyloplasts are starch-storing plastids
: Unpigmented plastids that resemble proplastids but contain strach granules.
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S: starch granules TEM of amyloplasts containing many starch granules (S)
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3) Several categories of plastid are named for their color
- Leucoplast are colorless plastids involved in the synthesis of monoterpenes, the volatile compounds
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TEM showing leucoplasts (L) in
an active secreting glandular trichome of pepperment.
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Etioplast TEM of an etioplast PR: large prolamellar body
T: associated unstacked thylakoids EN: envelope membrane
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An early stage of grana thylakoid development in a greening etioplast
PR An early stage of grana thylakoid development in a greening etioplast PR: prolamellar body
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Chloroplasts GT: grana thylakoids ST: stroma thylakoids
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Chromoplast (yellow, orange, or red….)
TEM of chromoplast of a ripe fruit of Jerusalem cherry. The dense bodies in the plastid contain carotenoids (or xanthophylls)
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4) The outer and inner membrane of the plastid envelope
differ in composition, structure and transporter functions Rich in galactolipids, poor in phopholipids Out memebrane: nonspecific pore protein pass size : 10 kd Inner membrane: specific transporters pass small uncharged
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5) The photosynthetic grana and stroma thylakoid membranes
form a physically continuous three-demensional network
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TEM depicting a single granum and associated stroma thylakoids
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The spatial relationship between stacked granna
and interconnecting stroma thylakoids
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TEM revealing difference in grana and stroma thylakoids
PS II PS I and ATP synthase TEM revealing difference in grana and stroma thylakoids
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6) Plastids are partially autonomous, encoding and
synthesizing some of their own proteins
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7) Plastids reproduced by division of existing plastids
TEM of dividing etioplast
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8) Plastids are inherited maternally in most flowering plants
but paternally in gymnosperms
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9) Plastids synthesize chlorophylls, cartenoids
and fatty acids and reduce some inorganic nutrients
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Mitochondria : contain the respiratory machinery that generates ATP
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TEM of a mitochondrion in a bean root tip cell
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Similarity in the basic architecture of all mitochondria reflects
the university of their mechanism for generating energy
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An unusual phopholipid component of inner mitochondrial membrane
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TEM depicting a longitudinal section through a transfer cell mitochondria
Arrows indicate ribosomes
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2) Small solutes cross the outer and inner mitochondrial membranes sequencially,
whereas large proteins destined for matrix cross both membrane simultaneously
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3) Mitochondria resemble prokaryotes in numerous important properties.
Mitochondria possess the genetic capacity to make some of their own proteins. Mitochondrial ribosomes resemble those of prokaryotes. Mitochondria reproduce by fission.
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TEM of a mitochondrion in a bean root tip cell,
showing the final phase of division
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4) Like plastids, mitochondria are semiautonomous and possess
the genetic machinery to make some of their own proteins Distinctive features of plant mitochondria genome 1. large size and complexity (about 10 fold larger than animal ones) 2. Genome: do not contain a set of tRNA genes (16 tRNAs specific to amino acids) 3. Contains some chloroplast DNA sequences (tRNA genes, but not functional)
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