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Chordata
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Phylum Chordata Bilateral, Deuterostomate development Notochord
Dorsal hollow nerve cord Pharyngeal slits Muscular Post-anal tail Segmented musculature Repeating units called somites
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Chordate features Oldest group ( ancestral) Urochordata
The Tunicates, Sea Squirts Chordate Features found in larval phase Aid in dispersal, adults are sessile. Today’s sessile tunicates are derived trait
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Chordate Phylogeny
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Subphylum Urochordata
Only larva has chordate characteristics #63-x-section #65
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Tunicates
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Tunicate larvae
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Tunicates
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atrial opening (water that passed through pharynx leaves this way)
nerve cord notochord gut oral opening atrial opening (water that passed through pharynx leaves this way) pharynx with gill slits Fig. 24.3, p. 385
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Subphylum Cephalochordata
Come about by Paedogenesis (?) Precocious sexual maturity in larvae Adults now have all the chordate traits, and are motile The lancelets have only a slight swelling , “anterior ganglia’ brain?
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Subphylum Cephalochordata
# 66
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Amphioxus
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Chordate Phylogeny
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Subphylum Craniata Cephalized Chordates Two sets of Hox genes
Brain, eyes, etc. Skull Two sets of Hox genes Neural Crest – Infolding of ectoderm Cells spread through developing body Form neurons and other features Teeth, facial bones, Pharanygeal slits paired with muscles & nerves that pump water through slits More active metabolism
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Class Mixini Jawless craniate Mixini – the hagfishes (not a fish)
Have cartilagnous, skull and notochord
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Super Class - Vertebrata
More extensive skull Backbone composed of vertebrae Originally prongs of cartilage dorsally along nototchord protecting nerve chord Later took over mechanical role of notochord Later fins and other appendages form along vertebrae
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Chordate Phylogeny
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Class Cephalaspidomorphi
Lampreys Have cartilaginous vertebrae-like extensions along notochord Still jawless
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Gnathostomes Vertebrates with true jaws Additional Hox gene cluster
Larger brains, better sense of smell sight Lateral line system to sense water movement Mineralized endoskeleton Two sets of paired appendages. these paired appendages first functioned in swimming. In tetrapods, the appendages are modified as legs on land.
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Class Chondrichthyes
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What’s New in Bony Fish Bony Skeleton
Single Gill Opening – Operculum bellows water over gills Swim bladder – gas from blood fills bladder, released to control buoyancy
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Swim Bladder Muscular Valve Gas Gland
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Lobed-finned fish vs. Amphibian Bones
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Class Amphibia
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REPTILIA - Amniotes
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MAMMALS
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Monotremes Warm blooded Have hair Lay eggs
Young hatch and live outside mother Make milk in glands, no nipples
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Marsupials Live birth to underdeveloped young. Placenta forms, but not a long a time. Young crawl to pouch Physically attach to nipple in pouch and feed off milk, finish development while nursing. Stay with mother in pouch until able to survive outside
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Eutherian mammals “Placental mammals”- live birth
Young held inside past egg feed development feed trough an umbilical attachment to the placental Born more developed than marsupials Feed off milk from breast- Not physically attached to nipple
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