1. Basal Tricolpates Finally, true dicots

2. Basal Angiosperms, view 1

3. Basal Angiosperms, view 2

4. Characters of Tricolpates Tricolpate pollen Floral parts whorled, members alternating Staminal filaments slender Sieve element plastids with starch grains Nuclear & chloroplast sequence data

5. Basal Tricolpates (true dicots)

6. Basal Tricolpates – Utah View Ranunculales, order of buttercup family, is important Ranunculaceae, Berberidaceae, Papaveracae Platanales of some interest Proteales – fascinating but mostly southern hemisphere

7. Ranunculales Mostly herbaceous Toothed to lobed or compound leaves Floral parts distinct, calyx, often many stamens, ovary superior, seeds with tiny embryos and much endosperm Alkaloid berberine Nuclear and chloroplast DNA

8. Papaveraceae (inc. Fumariaceae) Deciduous sepals Carpels 2, connate Fruit a capsule Nearly all have laticifers Also has properties of Ranunculales

9. Papaveraceae (2) Herbs or shrubs Laticifers present (opium), alkaloids Lvs simple but lobed or dissected, pinnate venation Flowers bisex, radial or bilat, Sepals 2, deciduous, petals 4 or 6 (often crumpled), sometimes united distally, stamens many to 2*3, carpels 2, united, superior ovary, placentation parietal, fruit a capsule

10. Dendromecon Argemone Dicentra Corydalis Lump or split?

11. A family of exceptions Mostly herbaceous, some shrubs, vines Lvs usually alt or spiral, usually simple, oft lobed or dissected, usually pinnate venation, usually no stipules Flowers usually bisex, usually radial, usually not 3- merous, tepals distinct, calyx usually distinct; stamens usually many, carpels usually 5+, US distinct, ovary superior Fruit follicles (dehiscent, >1 seed) or achenes (indehiscent, 1 seed) Ranunculaceae

12. Many stamens No hypanthium No stipules Ranunculaceae

13. Berberidaceae Herbs or shrubs (ours), yellow wood (berberine) Lv usually alt to spiral, simple to compound, entire to serrate or spinose-serrate Flrs bisex, rad, usually 3 merous, calyx distinct, inner petals (staminodes?) nectar- producing, stamens 4-many,anthers opening by flaps, carpel 1, ovary superior; fruit usually berry

15. Platanaceae Trees, monoecious Branched hairs Lvs alt or spir, usually palmate, gland tooth, petiole covers axil bud, stipules Flr unisex, reduced. Sep 3-7,pet 3-7, st 3-7, carp 5-9, distinct, ov sup. Fruit achene, with bristles

16. www.hope.edu/.../BasalEudicots/ Platanaceae web1.p15132387.pureserver.info/.../ Platane/ Platanaceae