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ORIGIN OF SPECIES CH 24
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Speciation: origin of new species Microevolution: changes in allele frequencies Macroevolution: changes that result in formation of different species
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I. The Biological Species Concept Biological species: group of populations whose members can breed and produce viable, fertile offspring
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A. Reproductive Isolation biological barriers that prevent formation of viable fertile offspring Can be prezygotic or postzygotic
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Prezygotic barriers:Prevent the formation of a zygote (fertilized egg) – Habitat Isolation: if 2 populations occupy different habitats, they don’t encounter each other to mate – Behavioral Isolation: Differences in behavior prevent mating – Temporal Isolation: Different breeding times prevent mating – Mechanical Isolation: Genitalia are incompatible – Gamete isolation: sperm doesn’t fertilize egg
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Postzygotic Barriers: Prevents the formation of normal fertile offspring – Embryo formed is not viable – Offspring produced are weak and die – Offspring produced are sterile
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II. Mechanisms of Speciation
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Allopatric Speciation – Results from lack of gene flow between populations that have been geographically isolated – Populations evolve independently due to mutation, natural selection, genetic drift – Reproductive isolation can result from genetic divergence
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Sympatric speciation – Evolution of different species WITHOUT geographic isolation – Polyploidy in plants is the doubling of chromosomes from one generation to the next resulting in reproductive isolation – Appearance of new ecological niches in an area can result in sympatric speciation
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Polyploidy
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III. Hybrid Zones Regions where two species with incomplete reproductive barriers meet and produce hybrid offspring Usually a band where 2 species meet
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3 possible outcomes in the hybrid zone: – Reinforcement of reproductive barriers: hybrid offspring are less fit so rate of hybridization decreases – Fusion: if hybrids are as fit as parents there could be enuf mating and gene flow between the populations that they fuse into 1 species – Stability: extensive gene flow prevents the continued selection of reproductive barriers
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