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Curvilinear isoclines might be more realistic

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Presentation on theme: "Curvilinear isoclines might be more realistic"— Presentation transcript:

1 Curvilinear isoclines might be more realistic

2 Mutualism Equations (pp. 234-235, Chapter 11)
dN1 /dt = r1 N1 ({X1 – N1 + a12 N2 }/X1) dN2 /dt = r2 N2 ({X2 – N2 + a21 N1 }/X2) (X1 – N1 + a12 N2 )/X1 = 0 when N1 = X1 + a12 N (X2 – N2 + a21 N1 )/X2 = 0 when N2 = X2 + a21 N1 If X1 and X2 are positive and a12 and a21 are chosen so that isoclines cross, there is a stable joint equilibrium Intraspecific self damping must be stronger than interspecific positive mutualistic effects.

3 N2* N1* Note: when together, both species reach higher equilbrium population density

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6 Evidence of Competition in Nature. often circumstantial. 1
Evidence of Competition in Nature often circumstantial 1.Resource partitioning among closely-related sympatric congeneric species (food, place, and time niches) Complementarity of niche dimensions 2. Character displacement 3. Incomplete biotas: niche shifts 4. Taxonomic composition of communities

7 Major Foods (Percentages) of Eight Species of Cone Shells, Conus,
on Subtidal Reefs in Hawaii __________________________________________________________ Gastro- Entero Tere- Other Species pods pneusts Nereids Eunicea belids Polychaetes flavidus lividus pennaceus 100 abbreviatus ebraeus sponsalis rattus imperialis

8 MacArthur’s Warblers (Dendroica)
Robert H. MacArthur

9 John Terborgh

10 John Terborgh

11 Time of Activity Seasonal changes in activity times

12 Complementarity of Niche Dimensions, page 276
Anolis Thomas W. Schoener

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14 Prey size versus predator size

15 Peter R. Grant David Lack

16 Character Displacement in Hydrobia mud snails in Denmark
Snail shell length, mm

17 Hutchinsonian ratios among short wing Accipiter hawks
Thomas W. Schoener

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21 Hutchinsonian ratios among Australian Varanus lizards


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