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Curvilinear isoclines might be more realistic
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Mutualism Equations (pp. 234-235, Chapter 11)
dN1 /dt = r1 N1 ({X1 – N1 + a12 N2 }/X1) dN2 /dt = r2 N2 ({X2 – N2 + a21 N1 }/X2) (X1 – N1 + a12 N2 )/X1 = 0 when N1 = X1 + a12 N (X2 – N2 + a21 N1 )/X2 = 0 when N2 = X2 + a21 N1 If X1 and X2 are positive and a12 and a21 are chosen so that isoclines cross, there is a stable joint equilibrium Intraspecific self damping must be stronger than interspecific positive mutualistic effects.
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N2* N1* Note: when together, both species reach higher equilbrium population density
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Evidence of Competition in Nature. often circumstantial. 1
Evidence of Competition in Nature often circumstantial 1.Resource partitioning among closely-related sympatric congeneric species (food, place, and time niches) Complementarity of niche dimensions 2. Character displacement 3. Incomplete biotas: niche shifts 4. Taxonomic composition of communities
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Major Foods (Percentages) of Eight Species of Cone Shells, Conus,
on Subtidal Reefs in Hawaii __________________________________________________________ Gastro- Entero Tere- Other Species pods pneusts Nereids Eunicea belids Polychaetes flavidus lividus pennaceus 100 abbreviatus ebraeus sponsalis rattus imperialis
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MacArthur’s Warblers (Dendroica)
Robert H. MacArthur
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John Terborgh
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John Terborgh
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Time of Activity Seasonal changes in activity times
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Complementarity of Niche Dimensions, page 276
Anolis Thomas W. Schoener
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Prey size versus predator size
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Peter R. Grant David Lack
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Character Displacement in Hydrobia mud snails in Denmark
Snail shell length, mm
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Hutchinsonian ratios among short wing Accipiter hawks
Thomas W. Schoener
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Hutchinsonian ratios among Australian Varanus lizards
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